Natalie Wolchover assembles mounting evidence that animals have morals:
Much of [Marc] Bekoff's research has focused on wolves and coyotes — both of which live in tight-knit groups governed by strict rules. … Breaking these rules of engagement — or other rules, such as taking more than one's fair share of food — is serious business among wolves and coyotes. "There is a consequence of being labeled a cheater," Bekoff said. Others stop bonding with the "immoral" pack member, and eventually it wanders away from the group, usually resulting in an early death because it no longer receives the benefits of pack living.
Previous coverage of Bekoff's work here.
The Dish 